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Before leaving the store and locking the doors behind them, they turn on the alarm, and check if other employees took care of their duties (such as swiping the floor or restocking the shelves).
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They typically come first in the morning and leave last in the evening. They should come to the store before everyone else, open the doors, turn off the alarm, and check if everything is in place, and the store is ready to open for customers. In most retail stores, key holder is simply a shift supervisor with a few extra responsibilities.